Spigots: AC = aciniform gland spigots(s), CO = colulus
, CY = cylindriform gland spigot(s), PI = pyriform gland spigot(s), SH = serrate hairs.
Of the similar genera (Table 1), the new genus Himalmartensus is similar to Rubrius and some Macrobunus species by having a single hairy colulus
The link between maternal care and subsociality led to a prediction: because maternal care in theridiids optimized to the "lost colulus
clade" (Anelosimus plus Theridiinae, see Agnarsson 2004 fig.
This evidence is consistent with predictions based on phylogenetic analysis that maternal care is primitively present in the lost colulus
clade, the lineage containing all social theridiids.
Spinnerets with colulus
represented by two setae; ALS conical, 2-segmented, contiguous, twice the width of the PLS; ALS with 3 spigots (piriform) and 3 nubbins; PMS with 3 spigots (aciniform) and 2 nubbins; PLS with 3 spigots (aciniform) and 3 nubbins.