More generally, by concentrating on towns the paper misses the opportunity to point to the fact that instead of the often assumed urban-rural dichotomy, which the paper rightly points out is misleading, there is a polychotomy encompassing urban-rural and central-remote locations.
Nevertheless, the ospAB phylogeny contains 1 polychotomy. Concatenating the data sets for ospA and ospB sequences is appropriate because these loci lie contiguously in a single operon, and little evidence exists for recombination either within or between these genes (see below).
Some minor disagreements have occurred, as in Theriot's (1988) criticism of Sluiman's (1985) failure to resolve grouping relationships of Coleochaete, other charophytes, bryophytes and tracheophyes (all arising in Sluiman's work as a sort of polychotomy).
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