1997) found a sharply toothed premaxilla
, mandible, vomer, and palatine, as well as fine-toothed gill arches and pharyngeal jaws.
Outer tooth row of premaxilla
with about 22 teeth, inner row with somewhat fewer teeth.
The maxillary segment contains its anterior articulation with the prenasal, dorsal articulation with the lacrimal, and ventral articulation with the premaxilla
75 in SL; dorsal at midbody; trunk scales 5/38/3', 12 predorsal, 14 around caudal peduncle; base of caudal covered with small scales extending onto the proximal 1/4 or 1/3 of the middle of the lobes, not along their border; anal fin iii,18-19, no hooklets observed; S03 entire, except a small "naked" triangle dorso-posteriorally; premaxilla
with 4 tricuspid teeth in the outer row and 4 pentacuspid ones in the inner row, all teeth usually black, the cuspids long; maxilla short, just reaching level of anterior margin of eye, with one broad black tooth; mandible 4 pen-tacuspid followed by about 8 much smaller tricuspid teeth.
and dentary with many irregularly distributed unicuspid, slightly curved teeth of different size, a small number of larger ones on the outer row of upper and lower jaw.
Five skull bones were chosen to obtain measurements for the relationships: the dentary, maxilla, premaxilla
, opercle, and cleithrum (Fig.
Jaws about equal, oblique, premaxilla
more or less straight, without abrupt bend between anterior horizontal portion and lateral part; maxilla ends below anterior edge of eye or slightly anterior to this point; maxillary length 3.
By 10 mm BL, premaxilla
and dentary bones were covered with caniniform teeth.
with outer row of ~20 small, slender teeth with tips curved posteriorly and visible below upper lip when mouth closed.
Ascending process of premaxilla
horizontal, its distal end almost above center of eye.
2% SL; teeth in upper jaw in 2-3 rows anteriorly, becoming two rows about midway along premaxilla
, ending in single row at posterior end of premaxilla
; all teeth conical with slightly recurved tips; teeth in outer row longest; teeth in lower jaw in two rows anteriorly, becoming single row about half way along dentary; teeth conical with slightly recurved tips; single recurved canine located at mid-dentary; tongue slightly emarginate; eye large, 10.
Small villiform teeth are present along the premaxilla