Telia generally abaxial, rarely amphigenous, on leaves and petioles, globose, blackish brown, scattered, pulverulent, 0.5-1.6 x 0.4-0.6 um.
UREDINIA amphigenous, mostly hypophyllous, cinnamon brown, and scattered, rounded, 0.3-0.7 mm diameter, pulverulent.
Probasidia cylindrical to clavate, becoming bifurcate, 4050 x 56 m.
Uredinia were epiphyllous, amphigenous
to mostly adaxial and yellowish orange to mostly cinnamon.
Telia amphigenous, mainly hypophyllous, subepidermalbecoming erumpent, arranged in circles and associated with leaf spots up to 8 mm diameter, 150-300 pm diameter, cupulate, peridium 75-150 pm above leaf surface.
Spermogonia amphigenous, when hypophyllous irregularly scattered among aecia.
, subcircular to angular-irregular, 1-5 mm diam., at first dark brown to almost black, later center paler, medium brown, dingy brown to grayish-brown, margin indefinite or a pale center surrounded by a narrow to wide dark border, occasionally with a narrow diffuse yellowish halo.
Basidiomata up to 6 X 3 cm, branched, gregarious; stipe up to 3 X 1.2 cm, irregular in cross-section, usually matted with hyphal layer identical to subiculum; branches in 1-5 ranks, irregularly dichotomous, ascending, flexuous, often covered with subicular-like layer downward, gray-drab, tandrab to pinkish tan ("avellaneous" "light pinkish cinnamon," "fawn color;" 7A2, 7B3, 7C5); hymenium amphigenous
above, often clearly unilateral below; internodes diminishing irregularly upward; axils narrowly rounded.
Uredinia amphigenous but mostly hypophyllous, intermixed, scattered, roundish, pulverulent, blackish brown, 0.14-0.3 x 0.2-0.3 mm.
Telia amphigenous but mostly on abaxial surface, black, compact, sub-epidermal, 0.04[?]0.07 x 0.06[?]0.14 mm.
Uredinia amphigenous, subepidermal, golden brown to blackish brown, present in rows, scattered but somewhat compact, naked, 0.06-0.11 x 0.094-2 mm.
Uredinia amphigenous, scattered, cinnamon brown, covered by ruptured epidermis, naked, 0.07-0.1 x 0.09-0.2 mm.