Bosch, "Advertisement calls of the midwife toads Alytes (Amphibia, Anura, Discoglossidae
) in continental Spain," Journal of Zoological Systematics and Evolutionary Research, vol.
The Lano association comprises at least five taxa: one indeterminate albanerpetontid Allocaudata (represented by incomplete dentaries, axis, and fragmentary humeri), one indeterminate Caudata (atlas, trunk vertebrae, humerus), and at least three anurans: one indeterminate Discoglossidae
(maxillaries, angulars, vertebrae, urostyle, humeri, and ilia), one indeterminate Palaeobatrachidae (angulars, presacral vertebra, synsacrums, humeri, and ilia), and at least one indeterminate anuran (maxillaries, humeri and ilia that could belong to one or several species) (Duffaud and Rage, 1999; Duffaud, 2000).
(2003) also reported isolated ilia from the Early Cretaceous of Morocco as belonging to the Laurasian clade Discoglossidae
. However, these specimens are very poorly preserved, and the combination of traits exhibited by these remains (e.g., posterodorsally extended supra-acetabular expansion, morphology of dorsal tuberosity) is not exclusive of discoglossids, being also present in calyptocephalellid and ceratophryid anurans.