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The fossil record shows that some Devonian fossil plants (genera Aglaophyton, Rhynia and Asteroxylon, 400 Myr) already had vesiclelike structures similar to those of Glomus
The telome theory is psilopsidcentered, for the Upper Silurian to Mid-Devonian Psilophytales (Hicklingia, Taenocrada, Zosterophyllum, Rhynia, Horneophyton, Psilophyton, etc.