Description: Herbs perennial or annual, tufted; culms scapose or few noded; leaves eligulate; inflorescence paniculate, with few to many spikelets, lower primary bracts leaflike, sheathing; spikelets with 3-6 persistent glumes, of increasing length, flowers bisexual, bristles spinulose to plumose, anthers apiculate
and conspicuously greenish yellow, achene obovate to oblong, beaked, surface smooth to reticulated, embryo more or less rhomboid to top-shaped with a tapered scutellum, notch present below root cap (embryo of the Carpha-type).
, Debaryomyces hansenii , Kloeckera apiculate
, Candida membranifaciens , and so forth.
+ (present);--(absent); Ob (obtuse); rd (round); Ac (acute); Ov (ovate); api (apiculate
); El (elliptic); Obv (obovate); An (antrorse); pt (patent); rt (retrorse).
The species of non-Saccharomyces yeast that are naturally present on grapes at the highest numbers belong mainly to the apiculate
yeast genus Hanseniaspora (mainly Hanseniaspora uvarum teleomorph and Kloeckera apiculata) and other species of the genera Candida, Pichia, Kluyveromyces and Metschnikowia.
turris in valve morphology and size but differs in the apiculate
shape of the apices (Metzeltin & Lange-Bertalot, 1998).
Rhizomes long creeping, covered with membranous ovate-apiculate scales; rhizome scales comose in distinct grades on dorsal surface, bicolorous with a dark central streak of sclerotic cells and pale margins, which are denticulate to erose; blade lanceolate to oblanceolate, basally pinnatisect and lobed to pinnatifid distally, with proximal lobe free; laminar scales scattered on abaxial surface and glabrescent on adaxial surface, with round to ovate scales with apiculate
or caudate apices, center obscure brown, margins pale brown, fimbriate to ciliate; midribs and secondary veins not visible; sori oblong, with orbicular, peltate scales similar to those in the lamina; spores bilateral, abortive, prominently verrucate.
Leaflets obovate to elliptic, 3-18 mm long, 3-9 mm wide, pubescent, upper half serrate, apiculate
. Inflorescence axillary, racemes pedunculate, peduncle 2-4 cm long, compact, 10-50 flowers.
Terminal, hairs straight, slightly curved or wavy ; Ascospores smaller to medium, size ranges between 9-13 x 6-9.5 [micro]m, apiculate
Five years ago, it documented eight species-"pipisik puti" (Avicennia marinae), "pipisik pula" (Avicennia officinalis), "pagatpat" (Sonneratia alba), "bakawan lalaki" (Rhizophora apiculate
), "baka-wan babae" (Rhizophora mucronata), "giting-giting" (Acanthus ilicifolius), "buta-buta" or "lipata" (Excoecaria agallocha) and "nipa" or "sasa" (Nypa fruticans).
Role of Apiculate
Yeasts on Organoleptic Characteristics of Wine.