The primary parietal cells divided repeatedly through periclinal and anticlinal divisions to form the subepidermal endothecium
and two middle layers (Fig.
The young anther wall (see Figure 2E), when microspore mother cells are present, consists of a uniseriate epidermis, an endothecium
formed by thin cell walls, one middle layer of cells similar to endothecium
cells, and one layer of secretory tapetai cells.
From the cross section we see that the anther wall consists of layers from outer to inner: epidermis, endothecium
, middle layer and layer of tapetum (Fig.
When orbicules are absent, the locule wall is smooth and endothecium
thickenings may be pronounced (Fig.
These crystals were also observed in the sepals, petals, placenta, style, anther connective and endothecium
. The clawed petals are light pink as is characteristic in the genus (Fig.
The microscopic observation of the flowers showed straight hairs, anther endothecium
, striate pollen grains, and parts of epidermis containing oxalate druses (Figure 12).
Immunolocalization of CP4 EPSPS in anther sections revealed little to no accumulation in the tapetum that nourishes the developing pollen and the microspore mother cells that are antecedents to pollen, despite presence of CP4 EPSPS in surrounding anther tissue (endothecium
, epidermis and connective/vascular tissue) (Fig.
2, 3(I-II), the microsporangium wall of sterile anthers consisted of epidermis, endothecium
, middle layer, and tapetum.
Then the wall differentiated to epidermis, endothecium
, middle layer and tapetum layer (Fig1 L).
The anther is formed by a unistratified epidermis, and the endothecium
is composed of cells, with elongated periclinal cells and irregular wall thickness (Figure 3e-f).
The main characters to justify the transference are: the short apical anther appendage; the glandular hairs in corolla, leaves, and fruits; the pollen morphology; the welldeveloped stylopodium; the widened style; and the endothecium
Each microsporangium displays four wall layers (from outside in: epidermis, endothecium
, middle layer, and parietal layer), a secretory tapetum and the central male cells.