Tree 10-15 m tall, small young branches yellowish gray to purplish brown, mature branches yellowish gray to yellowish brown, Stipule caduceus, linear lanceolate, apex acuminate, young petiole densely hairy, soon hairless; leaf blade ovate to broadly ovate, glabrous or thickly hairy when young, soon glabrescent
, base rounded or sub-cordate or sometime cordate, lustrous green above, margin long spinulose serrate, apex shortly acuminate or long tapering acute apex.
Rhizomes long creeping, covered with membranous ovate-apiculate scales; rhizome scales comose in distinct grades on dorsal surface, bicolorous with a dark central streak of sclerotic cells and pale margins, which are denticulate to erose; blade lanceolate to oblanceolate, basally pinnatisect and lobed to pinnatifid distally, with proximal lobe free; laminar scales scattered on abaxial surface and glabrescent
on adaxial surface, with round to ovate scales with apiculate or caudate apices, center obscure brown, margins pale brown, fimbriate to ciliate; midribs and secondary veins not visible; sori oblong, with orbicular, peltate scales similar to those in the lamina; spores bilateral, abortive, prominently verrucate.
with tector and dendroid trichomes on the abaxial surface; calyx 5-lobed, glabrous, with nectaries; corolla magenta with yellow nectar guides on the fauces ...
alpestris Boiss, with denticulate to serrate leaves, mucronate teeth, below glabrescent
to pubescent, petiole up to 0.9 cm, with sinuate secondary ribs and abundant fasciculate, multiradiate and stellate trichomes; later named by COUTINHO (1888) as: Q.
Leaves widely obovate to widely depressed obovate, widely elliptic to oblate, widely ovate-circular, very widely ovate to widely depressed ovate, length/width ratios 1.2-0.8:1; apices commonly retuse, sometimes rounded; crenate-dentate; leaves when yet unfolded adaxially subglabrous, with minute glandular hairs throughout, pubescent and tomentose over teeth, and puberulent here and there on surface, pubescent from a few teeth apices, tufted on midvein and some secondary veins, when newly unfolded adaxially subglabrate except at apex and base; abaxially pubescent (rays planar), golden-tomentellous, later glabrate, glabrescent
, gray trichomes, with primary to fifth order veins raised; white striae (wax?) along veins.
Leaves 12 to 14 in number, suberect-recurved, forming a broadly crateriform rosette; sheaths broadly elliptic, 19-25 x 17-20 cm, subdensely pale-brown lepidote on both sides, greenish or sometimes reddish near the apex; blades linear, not narrowed toward the base, 30-44 x 11-14 cm, glabrescent
abaxially, inconspicuously and sparsely white lepidote adaxially, rose-red throughout, more so adaxially or green toward the base and reddish near the apex, bearing dark green, sparsely arranged irregular spots, apex broadly acute to rounded and apiculate, apiculous ca.
Leaves rosulate; sessile to pseudopetiolate; blades entire to lyrate, lanceolate, elliptical, obovate to spathulate, pinnately veined, margin entire to runcinate, glabrous to glabrescent
above, densely pubescent beneath.
Shrubs 1-5 m tall, stems green, glabrous or glabrescent
when young, pale gray and glabrous and minutely lenticellate at maturity.
13 x 9.7 cm, densely brown lepidote on both sides, dark castaneous toward the base, coarsely spinose at the apex; blades sublinear, slightly narrowed toward the base, suberect-arcuate, 22-30 x 4.7-5 cm, inconspicuously and sparsely white-lepidote near the apex to glabrescent
, green, lustrous mainly abaxially, apex acuminate and apiculate, margins densely and coarsely spinose at the base with spines 2-5 x 1.5-2 mm, narrowly triangular, dark castaneous, straight to retrorse-uncinate, 4-8 mm apart, margins toward the apex laxly spinose, spines triangular, prevailing retrorse-uncinate, 1-2 x 1 mm, 7-15 mm apart.