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The upward bending of a leaf or other plant part, resulting from greater growth of the lower side than of the upper side.

hy′po·nas′tic adj.
American Heritage® Dictionary of the English Language, Fifth Edition. Copyright © 2016 by Houghton Mifflin Harcourt Publishing Company. Published by Houghton Mifflin Harcourt Publishing Company. All rights reserved.
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In plants, pri-to-pre-miRNA conversion also requires interaction of an RNA-binding protein HYPONASTIC LEAVES 1 (HYL1) and a C2H2-zinc finger protein SERRATE in nucleus.
After creation of plant pri-miRNA, a complex consisting of Dicer Like 1 enzyme (DCL1), the double-stranded RNA-binding protein HYL1 (hyponastic leaves 1), and the zinc-finger protein SE (serrate), with an assistance of the nuclear cap-binding complex, cleaves pri-miRNA to pre-miRNA [22, 28].
Yuan, "Structure of Arabidopsis HYPONASTIC LEAVES1 and its molecular implications for miRNA processing," Structure, vol.