Adult females have larviform
bodies lacking wings, legs, and eyes, and never leave their host insect.
This is not surprising given the fact that both the larviform
females and the larvae are confined to the forest floor where they prey upon millipedes (as reviewed in Arnett and Thomas, 2001).
Because of the specific geographic range and the female's larviform
appearance, the species has also been referred to as the "Appalachian glow-worm" (Lloyd 1971; Branham & Wenzel 2003).
One interpretation of their presence in domatia is that they are simply parasites of a mutualistic interaction and that domatium "design" cannot exclude tiny larviform
or flat mites and still allow access to potential mutualists.
However, a higher male ratio and an increase in the percentage of deformed larviform
pupae were recorded on the artificial diet.