The same number of joints in the tarsi is a character generally common to very large groups of beetles, but in the Engidae, as Westwood has remarked, the number varies greatly; and the number likewise differs in the two sexes of the same species: again in fossorial hymenoptera, the manner of neuration
of the wings is a character of the highest importance, because common to large groups; but in certain genera the neuration
differs in the different species, and likewise in the two sexes of the same species.
From Anaea it differs in having the big head, long and sharply pointed palpi and long stout antennae scarcely thickened towards the point, which are so characteristic of Prepona" and later he added "In neuration
this remarkable insect agrees with Prepona Bsd., the first subcostal branches are free as in Prepona and not united to the costal nervure, as in Anaea Hbn." (ROTHSCHILD, 1896).In 1904 Hans FRUHSTORFER in his preliminary study on Prepona placed Anaeomorpha erroneously in a separate section on the basis of the red androconia and the other features described above.