Bush and Van Auken (1986) concluded that sugarberry is a sciophyte
. They reported that seedlings of sugarberry achieved maximum growth under conditions of low light.
(1991) mentioned that plants exhibit several kinds of adaptation to the prevailing irradiance, phenotypic adaptation (modulative and modificative) and genotypic adaptation (heliophyte and sciophyte
plants) and that these adaptations are not mutually exclusive, bit superimposed so that they permit fine adjustments that guarantee the greatest possible efficiency of radiant energy utilization (Larcher 1983).
After fragmentation, forest community phylogenetic structure changed distinctly from clustered to dispersed, which was related to the relatively drier conditions in the forest following fragmentation, resulting in an increase of phylogenetically remote heliophytes and the loss of more closely related sciophytes
from the forest (Liu & Zhu, 2014).