spikelet with lower lemma glabrous or glabrous on both sides of the midnerve and alternately pilose and glabrous in the remaining zones.
1 cm long, pubescent externally, anthers subsessile
45 Pedicel (0) present in some or all flowers, (1) absent or highly reduced (flower sessile or subsessile
Among them subsessile
glandular trichomes are most widespread in all taxa examined.
Anthers are poricidal and 5.66 [+ or -] 0.57 mm long (n = 296) (Figure 4B and 4), and the filaments are much shorter than the subsessile
Leaves alternate, spiral or distichous; petiolate or subsessile
, presence of pulvinus; extrafloral nectaries absent or present, convex, sessile or stipitate, located between the pairs of leaflets or on the petiole; leaflets 1-many pairs, papyraceous to coriaceous, elliptic to oblong, lanceolate to obovate, apex acuminate, rounded to mucronate, base oblique.
, pedicel < 1 mm long, axillary to the 3-4 most distal bracts.
1.94 0.16 10 Orthion subsessile
(Standl.) Steyerm & 4.59 0.37 Standl.
Umbels concave, flat or convex, with a variable number of rays, pedunculate, rarely subsessile
. Fruit (1.5-)2-4 mm long, ellipsoid to subglobose, with the spines on the secondary ridges shorter than the width of the mericarps.
Flowers closely disposed, initially subsessile
, then pedunculate; pedicles 1-2 mm long.
The petioles are very short (4-5 mm versus 10-80 mm in the other species) and the bracts and upper leaves are subsessile
, Aspilia pluriseta, Guizotia scabra, Spilanthes mauritiana, Macaranga kilimandscharica (stem and leaves), Coleus kilimandschari, and Rumex bequaertii.